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After the recent sequencing of their genomes, the cnidarian Nematostella vectensis and the placozoan Trichoplax adhaerens have become favorite models for studying tissue regeneration and the evolution of multicellularity. More ancient basal metazoan phyla include the poriferan and ctenophore, whose evolutionary relationship has been clarified recently.

Our in silico and PCR-based methods indicate that the HSPG2 gene is conserved in both the placozoan and cnidarian genomes, but not in those of the ctenophores and only partly in poriferan genomes. HSPG2 also is absent from published ctenophore and Capsaspora owczarzaki genomes. The gene in T. These genetic constituents are found in isolated genes of various poriferan species, indicating a possible intronic recombinatory mechanism for assembly of the HSPG2 gene.

These data indicate that the complex HSPG2 gene evolved concurrently in a common ancestor of placozoans, cnidarians and bilaterians, likely along with the development of differentiated cell types separated by acellular matrices, and is activated to reestablish these tissue borders during wound healing. The human core protein consists of 4, amino acids and is modified with 3—4 heparan or chondroitin sulfate chains before secretion into the extracellular space, requiring a costly metabolic investment by the cell manufacturing it.

Perlecan performs signaling, adhesive and extracellular scaffolding roles [ 3 ], such that mutation of the gene has pleiotropic effects; complete loss of the perlecan gene is lethal in humans and mice [ 4 ]. HSPG2 orthologues have been studied in vertebrates, arthropods, and nematodes [ 5 , 6 ]. A goal of this study was to use the publicly available genomes of the basal metazoans Amphimedon queenslandica [ 7 ] and other porifera, Trichoplax adhaerens [ 8 ] and the cnidarian Nematostella vectensis [ 9 ] as well as the ctenophore Mnemiopsis leidyi [ 10 ] and the choanoflagellate Monosiga brevicollis [ 11 ] to infer the evolutionary events that created this large, complex, highly conserved proteoglycan.

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Intellectual Property Office Applicacion Nc,. Speed measurement and positioning systems are key elements in railway operation as parts of the Automatic Train Control ATC and Automatic Train Protection ATP systems, since braking curves and maximum distance permissible between trains are calculated through train position and speed. Therefore, improvements in the measurement precision and quality will result in safer, faster and more efficient railway networks. Desirable characteristics of position and speed estimation systems include: reliability, high precision, accuracy and availability over different weather conditions, low latency, protection against electromagnetic interferences, high data-rate, low cost and ease of installation and maintenance.

Therefore several disparate technologies of measurement and integration are commonly used, leading to multisensor systems. The most frequent technologies used include balises, tachometers, track circuits, Doppler radar, inertial navigation systems, global navigation satellite systems, and artificial vision. Advantages and drawbacks of each may be balanced by using sensor fusion systems with or without map-matching strategies.

A known-type of speed estimator system is based on the delay in the pass through irregularities of the rail track. In this model two sensors are placed separated by a distance d, in the direction of motion, and convert rail irregularities into two signals xl and x2. Ideally x2 is only a delayed version of xl. This estimator system has been proposed for several types of sensors including optical, eddy current and vibration sensors. However, the practical installation and maintenance of a system with sensors in the bogies is complex.

The sensors are exposed to very hostile mechanical conditions, and carrying signals to a processing unit presents several technical problems.

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